Portal:Amphibians

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The Amphibian Portal

Anaschisma

Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods, excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds, and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.

The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.

The earliest amphibians evolved in the Devonian period from sarcopterygian fish with lungs and bony-limbed fins, features that were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (mammal predecessors). The origin of modern amphibians belonging to Lissamphibia, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. However the emerging consensus is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. (Full article...)

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Selected frog article

Allobates femoralis

The Aromobatidae are a family of frogs native to Central and South America. They are sometimes referred to as cryptic forest frogs or cryptic poison frogs. They are the sister taxon of the Dendrobatidae, the poison dart frogs, but are not as toxic as most dendrobatids are. (Full article...)

List of selected frog articles

Selected salamander article

Fossil of the salamander Karaurus sharovi a non-urodelan caudatan from the Middle-Late Jurassic belonging to the family Karauridae

The Caudata are a group of amphibians containing the extant salamanders (the order Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults.

Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. (Full article...)

List of selected salamander articles
Ambystomatidae Cryptobranchoidea Greater siren Salamanders in folklore Salamandroidea Sexual selection in amphibians Sirenidae Urodela

Did you know? – show different entries

... that the Pyrenean brook salamander sometimes lives entirely underground in caves?
... that Rhachitomi, a group of temnospondyl amphibians, was established as a clade in 2013?
... that the specific name of Tambachia trogallas, the type species of the trematopid temnospondyl Tambachia, refers to the Thuringian bratwurst that was frequently eaten by the describers of the species?
... that the female tree frog Aplastodiscus leucopygius inspects the underground nesting chamber prepared by the male before accepting his advances?
... that the Italian pool frog (pictured) has been introduced into the United Kingdom?

Selected amphibian type

Oscaecilia ochrocephala (Caeciliidae)

Caecilians (/sɪˈsɪliən/; New Latin for 'blind ones') are a group of limbless, vermiform (worm-shaped) or serpentine (snake-shaped) amphibians with small or sometimes nonexistent eyes. They mostly live hidden in soil or in streambeds, and this cryptic lifestyle renders caecilians among the least familiar amphibians. Modern caecilians live in the tropics of South and Central America, Africa, and southern Asia. Caecilians feed on small subterranean creatures such as earthworms. The body is cylindrical and often darkly coloured, and the skull is bullet-shaped and strongly built. Caecilian heads have several unique adaptations, including fused cranial and jaw bones, a two-part system of jaw muscles, and a chemosensory tentacle in front of the eye. The skin is slimy and bears ringlike markings or grooves and may contain scales.

Modern caecilians are a clade, the order Gymnophiona' /ˌdʒɪmnəˈfaɪənə/ (or Apoda /ˈæpədə/), one of the three living amphibian groups alongside Anura (frogs) and Urodela (salamanders). Gymnophiona is a crown group, encompassing all modern caecilians and all descendants of their last common ancestor. There are more than 220 living species of caecilian classified in 10 families. Gymnophionomorpha is a recently coined name for the corresponding total group which includes Gymnophiona as well as a few extinct stem-group caecilians (extinct amphibians whose closest living relatives are caecilians but are not descended from any caecilian). Some palaeontologists have used the name Gymnophiona for the total group and the old name Apoda for the crown group'. However, Apoda has other even older uses, including as the name of a genus of Butterfly making its use potentially confusing and best avoided. 'Gymnophiona' derives from the Greek words γυμνος / gymnos (Ancient Greek for 'naked') and οφις / ophis (Ancient Greek for 'snake'), as the caecilians were originally thought to be related to snakes and to lack scales.

The study of caecilian evolution is complicated by their poor fossil record and specialized anatomy. Genetic evidence and some anatomical details (such as pedicellate teeth) support the idea that frogs, salamanders, and caecilians (collectively known as lissamphibians) are each others' closest relatives. Frogs and salamanders show many similarities to dissorophoids, a group of extinct amphibians in the order Temnospondyli. Caecilians are more controversial; many studies extend dissorophoid ancestry to caecilians. Some studies have instead argued that caecilians descend from extinct lepospondyl or stereospondyl amphibians, contradicting evidence for lissamphibian monophyly (common ancestry). Rare fossils of early gymnophionans such as Eocaecilia and Funcusvermis have helped to test the various conflicting hypotheses for the relationships between caecilians and other living and extinct amphibians. (Full article...)

List of selected amphibian type articles
Frog Toad Salamander

Selected images

Image 1
Spanish painted frog
Photograph: Benny Trapp
The Spanish painted frog (Discoglossus jeanneae) is a species of frog in the family Alytidae. Endemic to Spain, it mostly lives in open areas, pine groves and shrublands. It feeds mostly on insects and worms.
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Image 2
Ecnomiohyla rabborum
Photograph: Brian Gratwicke
Rabbs' fringe-limbed treefrog (Ecnomiohyla rabborum) is a large species of frog originally found in the forest canopies of central Panama. Only discovered in 2005, the species is thought to be extinct in the wild; only one specimen, a male at the Atlanta Botanical Garden, survives.
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Image 3
Southern brown tree frog
Photograph: JJ Harrison
The southern brown tree frog (Litoria ewingii) is a species of tree frog native to Australia found in a wide range of habitats. Reaching 45 millimetres (1.8 in), this species is generally brown, but green and green-striped morphs are also recorded.
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Image 4
Vocal sac
Photo: Benny Trapp
An Italian tree frog (Hyla intermedia) with an inflated vocal sac. Vocal sacs, flexible membranes of skin possessed by most male frogs, are generally used to amplify mating or advertisement calls. However, some species in the family Rhinodermatidae may also use the sac to store or transport their tadpoles.
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Image 5
Raorchestes parvulus
Photograph credit: Rushenb
Raorchestes parvulus is a small bush frog native to tropical southeastern Asia. Although the reproductive strategy of this species has not been studied, members of the family typically make a foam nest while mating in vegetation overhanging a stream. They create the foam by beating their legs, and the eggs are deposited into the nest and covered with seminal fluid before the foam hardens into a protective casing. When the eggs hatch, the young push their way out of the nest and fall into the water below.

This picture shows a male R. parvulus frog, photographed in Phu Kradueng National Park, Thailand, calling by inflating its vocal sac.
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Image 6
Marsh frog
Photo: Marie-Lan Nguyen
The marsh frog (Pelophylax ridibundus) is a species of true frog and the largest frog native to Europe; females of this sexually dimorphic species may be up to 17 centimetres (6.7 in) long. The marsh frog feeds mainly on insects, but it also eats smaller amphibians, fish, and rodents.
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Image 7
Common toads in amplexus
Photo: Bernie Kohl
Two common toads (Bufo bufo) in amplexus, a form of pseudocopulation found in amphibians and limulids. A male common toad grasps a female with his front legs as part of the mating process. Common toads stay in amplexus for several days. As the female lays a long, double string of small black eggs, the male fertilises them with his sperm; the gelatinous egg strings, which may contain 3000 to 6000 eggs and be 3 to 4.5 metres (10 to 15 ft) in length, are later tangled in plant stalks.
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Image 8
Wallace's flying frog
Photograph credit: Rushenb
Wallace's flying frog (Rhacophorus nigropalmatus) is a moss frog found in tropical southeastern Asia. It is named after the British naturalist Alfred R. Wallace, who collected the first known specimen of the species. It lives almost exclusively in trees, and when threatened, or in search of prey, will leap from a branch and splay its four webbed feet; the membranes between its toes and the loose skin flaps on its sides catch the air as it falls, helping it to glide. This individual was photographed in Khao Sok National Park, Thailand.
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Image 9
Eastern newt
Photo credit: Patrick Coin
A terrestrial subadult Eastern newt or red eft (Notophthalmus viridescens). Salamanders of the family Salamandridae with aquatic adult stages are called newts. Some newts, including the Eastern newt, have a juvenile terrestrial stage called the eft. The red eft has bright aposematic coloration to warn predators of its highly toxic skin.
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Image 10
Albanian water frog
Photograph: Benny Trapp
The Albanian water frog (Pelophylax shqipericus) is a species of true frog in the family Ranidae. It is native to Albania and Montenegro, where it lives in aquatic environments. The frogs are medium-sized. Males sometimes bear a distinctive bright green stripe down the length of the backbone, but otherwise are green to light brown in overall colouring with large black or dark brown spots. Females are olive green or light brown in colour and also bear brown or black large spots. The species is endangered and known populations are currently in decline.
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Image 11
Blue Mountains Tree Frog
Photo: Benjamint444
The Blue Mountains Tree Frog (Litoria citropa) is a moderate-sized species of tree frog, up to about 60 mm (2.4 in) in length. It is native to coastal and highland areas of eastern Australia, especially in the Blue Mountains, hence its name.
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Image 12
Frogspawn
Photo credit: Tarquin
In the life cycle of a frog, a female lays her eggs in a shallow pond or creek, where they will be sheltered from the current and from predators. The eggs, known as frogspawn hatch into tadpoles. The tadpoles develop gradually into adolescent froglets and finally the froglet develops into an adult frog.
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Image 13
Red-eyed Tree Frog
Photo credit: Carey James Balboa
The Red-eyed Tree Frog (Agalychnis callidryas) is a small-sized tree frog, reaching lengths of about 5 to 7 centimeters (3 inches), native to Neotropical rainforests in Central America. They are not poisonous and rely on camouflage to protect them. During the day, they remain motionless with their colorful parts hidden. Thus, they appear almost completely green, and well hidden among the foliage.
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Image 14
Common frog
Photo: Richard Bartz
The common frog (Rana temporaria) is found throughout much of Europe. Adults have a body length of 6 to 9 cm (2.4 to 3.5 in) and vary in colour, with the ability to lighten and darken their skin to match their surroundings. They will feed on any invertebrate of a suitable size and, apart from the breeding season, live solitary lives.
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Image 15
White's Tree Frog
Photo credit: liquidGhoul
Native to Australia, White's Tree Frog grows up to 10 centimetres in length and is a popular household pet. In captivity, they have an average lifespan of 16 years. Its skin secretions contain caerins, a group of peptides with antibacterial and antiviral properties. Other peptides have been found to destroy HIV without harming healthy T-cells.
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Selected toad article

Couch's spadefoot toad or Couch's spadefoot (Scaphiopus couchii) is a species of North American spadefoot toad (family Scaphiopodidae). The specific epithet couchii is in honor of American naturalist Darius Nash Couch, who collected the first specimen while on a personal expedition to northern Mexico to collect plant, mineral, and animal specimens for the Smithsonian Institution. (Full article...)

List of selected toad articles

Selected caecilian article

Tapalcua (Dermophis mexicanus)

Dermophis is a genus of worm-like amphibians in the family Dermophiidae, the Neotropical and Tropical African caecilians. They are found in the Middle America between southern Mexico and northwestern Colombia. Common names Mexican caecilians or Neotropical caecilians are sometimes used for them. (Full article...)

List of selected caecilian articles